Dan. [ǭ] became [ō] in East Danian; [ǭ] > [ō]
According to the Comparative Tables, the primitive rounded long [ǭ] (as well as [ǭ] from primitive [ā]) developed into [ō] in the East Danian dialect, instead of becoming the diphthong [ua] as it did in Ossiriandic (PE19/25). In earlier notes associated with the tables, Tolkien described this as a general development in Danian instead of unique to East Danian (PE19/26). Since there are no examples of this change attested in The Etymologies, it is hard to guess what Tolkien ultimately decided.
References ✧ PE19/26
Phonetic Rule Elements
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